The remains of male mountain lion P-64 were discovered on Monday after researchers hiked to his last known GPS point and found his remains, the National Park Service announced Friday. 

The 4-year-old mountain lion survived the Woolsey fire, but appeared to have died a few weeks afterward. Although the cause of death is not known at this time, his paws were visibly burned and the California Department of Fish and Wildlife will conduct a necropsy to discover the reason for his death.

“P-64 was a fascinating cat to study because he crossed our notoriously deadly freeways dozens of times,” said Jeff Sikich, biologist for Santa Monica Mountains National Recreation Area. “It’s very unfortunate that he was seemingly so successful surviving in this fragmented landscape and then died in the aftermath of a devastating wildfire. It’s of particular interest that he chose to travel back through a fresh burn area rather than retreat through urbanized areas to escape the fire.”

The mountain lion was tracked in the Simi Hills, north of Oak Park, when the Woolsey fire broke out on November 8. According to the NPS, he traveled throughout the Simi Hills, covering several miles before hunkering down in a remote area. He was located alive on November 26 by a telemetry device in an unburned portion of the Simi Hills. Researchers found his body near his last known GPS location on Monday, and he appeared to have been dead for days.

Also known as "the Culvert Cat," he surprised researchers by using a storm drain multiple times to cross under the 101 Freeway near Liberty Canyon in Agoura Hills. During the nine months that he was tracked by NPS researchers, he crossed the 101 and 118 freeways a total of 41 times and is the suspected father of four female kittens who were born in May.

According to the NPS, of the 13 mountain lions that are tracked in the region, 11 were in or around the fire perimeter and nine have survived the fire and appear to be moving normally.